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Araneomorphae

From Wikipedia, the free encyclopedia

Araneomorph spiders
Temporal range: Triassic–present
Nephila inaurata (Nephilidae)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Suborder: Opisthothelae
Infraorder: Araneomorphae
Subdivisions
Diversity
95 families

The Araneomorphae (also called the Labidognatha or "true spiders"[1]) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their close kin), where they point straight down. Araneomorphs comprise the vast majority (about 93%[2]) of living spiders.

Distinguishing characteristics

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Most spider species are Araneomorphae, which have fangs that face towards each other, increasing the orientations that they can employ during prey-capture. They have fewer book lungs (when present) – usually one pair – and the females typically live one year.

The Mygalomorphae have fangs that face towards the ground, and which are parallel to the long axis of the spider's body, thus they have only one orientation they can employ during prey capture. They have two pairs of book lungs, and the females often live many years.[3]

Spiders included

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Almost all of the familiar spiders are included in the Araneomorphae group, one major exception being the tarantulas. There are a few other Mygalomorphae species that live around homes or gardens, but they typically are relatively small and not easily noticed.

The Araneomorphae, to the contrary, include the weavers of spiral webs; the cobweb spiders that live in the corners of rooms, and between windows and screens; the crab spiders that lurk on the surfaces of flowers in gardens; the jumping spiders that are visible hunting on surfaces; the wolf spiders that carpet hunting sites in sunny spots; and the large huntsman spiders.

Systematics

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In older schemes, the Araneomorphae were divided into two lineages, the Hypochilae (containing only the family Hypochilidae), and the Neocribellatae. The Neocribellatae were in turn divided into the Austrochiloidea, and the two series Haplogynae and Entelogynae, each containing several superfamilies. Molecular phylogenetic studies have shown that the haplogynes in particular are not a monophyletic group. A 2020 study suggested the relationships among the major groups were as shown in the following cladogram.[4]

Araneomorphae
Haplogynae

The blue bar to the right shows the former Haplogynae in the sense of Coddington (2005).[5]

Table of families

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Key
Genera 1 ≥2 ≥10 ≥100
Species 1–9 ≥10 ≥100 ≥1000
Araneomorphae families[notes 1]
Family Genera Species Common name Example
Agelenidae 97 1420 araneomorph funnel-web spiders Hobo spider (Eratigena agrestis)
Amaurobiidae 26 202 tangled nest spiders Callobius claustrarius
Anapidae 59 233 Holarchaea novaeseelandiae
Ancylometidae 1 11
Anyphaenidae 58 649 anyphaenid sac spiders Hibana velox (yellow ghost spider)
Araneidae 198 3144 orb-weaver spiders Zygiella x-notata
Archaeidae 6 93 pelican spiders Madagascarchaea gracilicollis
Archoleptonetidae 2 8 Archoleptoneta gertschi
Arkyidae 2 38
Austrochilidae 2 9 Tasmanian cave spider (Hickmania troglodytes)
Caponiidae 21 153 Diploglena capensis
Cheiracanthiidae 15 376 Cheiracanthium mildei'
Cicurinidae 4 172
Cithaeronidae 2 9
Clubionidae 18 667 sac spiders Clubiona trivialis
Corinnidae 76 876 dark sac spiders Castianeira sp.
Ctenidae 48 605 wandering spiders Phoneutria fera
Cyatholipidae 23 58
Cybaeidae 23 301 Cryphoeca silvicola
Cycloctenidae 8 80
Deinopidae 3 68 net-casting spiders Asianopis subrufa (rufous net-casting spider)
Desidae 63 323 intertidal spiders Phryganoporus candidus
Dictynidae 51 460 Nigma walckenaeri
Diguetidae 2 16 coneweb spiders
Dolomedidae 7 128
Drymusidae 2 19 false violin spiders
Dysderidae 24 653 woodlouse hunter spiders Woodlouse spider (Dysdera crocata)
Eresidae 9 106 velvet spiders Eresus sandaliatus
Filistatidae 18 192 crevice weavers Southern house spider (Kukulcania hibernalis)
Fonteferreidae 1 1
Gallieniellidae 5 41
Gnaphosidae 153 2479 flat-bellied ground spiders Drassodes cupreus
Gradungulidae 8 18 large-clawed spiders Progradungula carraiensis (Carrai cave spider)
Hahniidae 29 240 dwarf sheet spiders
Hersiliidae 16 187 tree trunk spiders Hersilia savignyi
Homalonychidae 1 2
Huttoniidae 1 1 Huttonia palpimanoides
Hypochilidae 2 33 lampshade spiders Hypochilus thorelli
Lamponidae 23 192 White-tailed spider (Lampona spp.)
Leptonetidae 22 397 Tooth Cave spider (Tayshaneta myopica)
Linyphiidae 640 4940 dwarf / money spiders Linyphia triangularis
Liocranidae 35 354 liocranid sac spiders
Lycosidae 135 2490 wolf spiders Lycosa tarantula
Macrobunidae 26 92
Malkaridae 13 57 shield spiders
Mecysmaucheniidae 7 25
Megadictynidae 2 2
Mimetidae 8 164 pirate spiders Oarces reticulatus
Miturgidae 33 191 long-legged sac spiders
Myrmecicultoridae 1 1
Mysmenidae 17 188 spurred orb-weavers
Nesticidae 16 292 cave cobweb spiders Nesticella marapu
Nicodamidae 7 27
Ochyroceratidae 9 184 midget ground weavers Theotima minutissima
Oecobiidae 7 129 disc web spiders Oecobius navus
Oonopidae 115 1962 dwarf hunting spiders Oonops domesticus
Orsolobidae 30 189
Oxyopidae 9 448 lynx spiders Peucetia viridans (green lynx spider)
Pacullidae 4 38
Palpimanidae 20 182 palp-footed spiders
Penestomidae 1 9
Periegopidae 1 3
Philodromidae 30 527 philodromid crab spiders Philodromus dispar
Pholcidae 97 2029 daddy long-legs spiders Pholcus phalangioides
Phrurolithidae 25 410
Physoglenidae 13 72
Phyxelididae 14 68
Pimoidae 2 87 Pimoa cthulhu
Pisauridae 45 236 nursery web spiders Pisaura mirabilis
Plectreuridae 2 32
Prodidomidae 24 195
Psechridae 2 62
Psilodercidae 11 224
Salticidae 689 6808 jumping spiders Zebra spider (Salticus scenicus)
Scytodidae 4 253 spitting spiders Scytodes thoracica
Segestriidae 5 181 tubeweb spiders Segestria florentina
Selenopidae 9 282 wall spiders Selenops radiatus
Senoculidae 1 31
Sicariidae 3 176 recluse spiders Brown recluse (Loxosceles reclusa)
Sparassidae 97 1519 huntsman spiders Delena cancerides (Avondale spider)
Stenochilidae 2 13
Stiphidiidae 20 125 Tartarus mullamullangensis
Symphytognathidae 10 104 dwarf orb-weavers Patu digua
Synaphridae 3 13
Synotaxidae 5 38
Telemidae 16 107 long-legged cave spiders
Tetrablemmidae 27 153 armored spiders
Tetragnathidae 45 989 long jawed orb-weavers Leucauge venusta (orchard spider)
Theridiidae 131 2583 cobweb spiders Redback spider (Latrodectus hasselti)
Theridiosomatidae 22 150 ray spiders Theridiosoma gemmosum
Thomisidae 170 2169 crab spiders Misumena vatia (goldenrod crab spider)
Titanoecidae 5 67 Goeldia obscura
Toxopidae 14 82
Trachelidae 29 300
Trachycosmidae 20 148
Trechaleidae 17 136
Trochanteriidae 6 52
Trogloraptoridae 1 1 Trogloraptor marchingtoni
Udubidae 6 57
Uloboridae 19 283 hackled orb-weavers Uloborus walckenaerius
Viridasiidae 3 14
Xenoctenidae 4 33
Zodariidae 90 1306 Zodarion germanicum
Zoropsidae 28 186 Zoropsis spinimana


Extinct families

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Notes

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  1. ^ Unless otherwise shown, currently accepted families and counts based on the World Spider Catalog version 25.5 as of 7 February 2025.[6] In the World Spider Catalog, "species" counts include subspecies. Assignment to sub- and infraorders based on Coddington (2005, p. 20) (when given there).

References

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Citations

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  1. ^ Coddington, J.A. (2005). "Phylogeny and Classification of Spiders" (PDF). In Ubick, D.; Paquin, P.; Cushing, P.E.; Roth, V. (eds.). Spiders of North America: an identification manual. American Arachnological Society. pp. 18–24. ISBN 978-0-9771439-0-0. Archived from the original (PDF) on 9 December 2008. Retrieved 12 October 2008.
  2. ^ Bern, Natural History Museum. "NMBE - World Spider Catalog". research.amnh.org. Retrieved 2023-06-07.
  3. ^ "ABOUT SPIDERS". www.publish.csiro.au. Retrieved 2022-01-07.
  4. ^ Magalhaes, Ivan L. F.; Azevedo, Guilherme H. F.; Michalik, Peter; Ramírez, Martín J. (February 2020). "The fossil record of spiders revisited: implications for calibrating trees and evidence for a major faunal turnover since the Mesozoic". Biological Reviews. 95 (1): 184–217. doi:10.1111/brv.12559. ISSN 1464-7931. PMID 31713947. S2CID 207937170.
  5. ^ Coddington, Jonathan A. (2005). "Phylogeny and classification of spiders" (PDF). In Ubick, D.; Paquin, P.; Cushing, P.E. & Roth, V. (eds.). Spiders of North America: an identification manual. American Arachnological Society. pp. 18–24. Retrieved 2015-09-24.
  6. ^ "Currently valid spider genera and species". World Spider Catalog. Natural History Museum Bern. Retrieved 2025-02-07. Currently valid spider genera and species
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